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In the New Testament angels appear frequently as the ministers of God and the agents of revelation; and Our Lord speaks of angels as fulfilling such functions, implying in one saying that they neither marry nor are given in marriage. Naturally angels are most prominent in the Apocalypse. The New Testament takes little interest in the idea of the angelic hierarchy, but there are traces of the doctrine. The distinction of good and bad angels is recognized; we have names, Gabriel, and the evil angels Abaddon or Apollyon, Beelzebub. and Satan; ranks are implied, archangels, principalities and powers, thrones and dominions. Angels occur in groups of four or seven. In Rev. i.-iii. we meet with the "Angels" of the Seven Churches of Asia Minor. These are probably guardian angels, standing to the churches in the same relation that the "princes" in Daniel stand to the nations; practically the "angels" are personifications of the churches. A less likely view is that the "angels" are the human representatives of the churches, the bishops or chief presbyters. There seems, however, no parallel to such a use of "angel," and it is doubtful whether the monarchical government of churches was fully developed when the Apocalypse was written.

Nor does the Bible give any formal account of the nature of angels. It is doubtful how far Ezekiel's account of the cherubim and Isaiah's account of the seraphim are to be taken as descriptions of actual beings; they are probably figurative, or else subjective visions. Angels are constantly spoken of as "men," and, including even the Angel of Yahweh, are spoken of as discharging the various functions of human life; they eat and drink, walk and speak. Putting aside the cherubim and seraphim, they are not spoken of as having wings. On the other hand they appear and vanish, exercise miraculous powers, and fly. Seeing that the anthropomorphic language used of the angels is similar to that used of God, the Scriptures would hardly seem to require a literal interpretation in either case. A special association is found, both in the Bible and elsewhere, between the angels and the heavenly bodies, and the elements or elemental forces, fire, water, &c. The angels are infinitely numerous.

While the scriptural statements imply a belief in the existence of spiritual beings intermediate between God and men, it is probable that many of the details may be regarded merely as symbolic imagery. In Scripture the function of the angel overshadows his personality; the stress is on their ministry; they appear in order to perform specific acts.

ANGELICO, FRA , Italian painter. Il Beato Fra Giovanni Angelico da Fiesole is the name given to a far-famed painter-friar of the Florentine state in the 15th century, the representative, beyond all other men, of pietistic painting. He is often, but not accurately, termed simply "Fiesole," which is merely the name of the town where he first took the vows; more often Fra Angelico. If we turn his compound designation into English, it runs thus--"the Beatified Friar John the Angelic of Fiesole." In his lifetime he was known no doubt simply as Fra Giovanni or Friar John; "The Angelic" is a laudatory term which was assigned to him at an early date,--we find it in use within thirty years after his death; and, at some period which is not defined in our authorities, he was beatified by due ecclesiastical process. His baptismal name was Guido, Giovanni being only his name in religion. He was born at Vicchio, in the Tuscan province of Mugello, of unknown but seemingly well-to-do parentage, in 1387 ; in 1407 he became a novice in the convent of S. Domenico at Fiesole, and in 1408 he took the vows and entered the Dominican order. Whether he had previously been a painter by profession is not certain, but may be pronounced probable. The painter named Lorenzo Monaco may have contributed to his art-training, and the influence of the Sienese school is discernible in his work.

According to all the accounts which have reached us, few men on whom the distinction of beatification has been conferred could have deserved it more nobly than Fra Giovanni. He led a holy and self-denying life, shunning all advancement, and was a brother to the poor; no man ever saw him angered. He painted with unceasing diligence, treating none but sacred subjects; he never retouched or altered his work, probably with a religious feeling that such as divine providence allowed the thing to come, such it should remain. He was wont to say that he who illustrates the acts of Christ should be with Christ. It is averred that he never handled a brush without fervent prayer and he wept when he painted a Crucifixion. The Last Judgment and the Annunciation were two of the subjects he most frequently treated.

We have spoken of Angelico's art as "pietistic"; this is in fact its predominant character. His visages have an air of rapt suavity, devotional fervency and beaming esoteric consciousness, which is intensely attractive to some minds and realizes beyond rivalry a particular ideal--that of ecclesiastical saintliness and detachment from secular fret and turmoil. It should not be denied that he did not always escape the pitfalls of such a method of treatment, the faces becoming sleek and prim, with a smirk of sexless religiosity which hardly eludes the artificial or even the hypocritical; on other minds, therefore, and these some of the most masculine and resolute, he produces little genuine impression. After allowing for this, Angelico should nevertheless be accepted beyond cavil as an exalted typical painter according to his own range of conceptions, consonant with his monastic calling, unsullied purity of life and exceeding devoutness. Exquisite as he is in his special mode of execution, he undoubtedly falls far short, not only of his great naturalist contemporaries such as Masaccio and Lippo Lippi, but even of so distant a precursor as Giotto, in all that pertains to bold or life-like invention of a subject or the realization of ordinary appearances, expressions and actions--the facts of nature, as distinguished from the aspirations or contemplations of the spirit. Technically speaking, he had much finish and harmony of composition and colour, without corresponding mastery of light and shade, and his knowledge of the human frame was restricted. The brilliancy and fair light scale of his tints is constantly remarkable, combined with a free use of gilding; this conduces materially to that celestial character which so pre-eminently distinguishes his pictured visions of the divine persons, the hierarchy of heaven and the glory of the redeemed.

ANGEL-LIGHTS, in architecture, the outer upper lights in a perpendicular window, next to the springing; probably a corruption of the word angle-lights, as they are nearly triangular.

ANGERS, a city of western France, capital of the department of Maine-et-Loire, 191 m. S.W. of Paris by the Western railway to Nantes. Pop. 73,585. It occupies rising ground on both banks of the Maine, which are united by three bridges. The surrounding district is famous for its flourishing nurseries and market gardens. Pierced with wide, straight streets, well provided with public gardens, and surrounded by ample, tree-lined boulevards, beyond which lie new suburbs, Angers is one of the pleasantest towns in France. Of its numerous medieval buildings the most important is the cathedral of St. Maurice, dating in the main from the 12th and 13th centuries. Between the two flanking towers of the west fa?ade, the spires of which are of the 16th century, rises a central tower of the same period. The most prominent feature of the fa?ade is the series of eight warriors carved on the base of this tower. The vaulting of the nave takes the form of a series of cupolas, and that of the choir and transept is similar. The chief treasures of the church are its rich stained glass and valuable tapestry . The bishop's palace which adjoins the cathedral contains a fine synodal hall of the 12th century. Of the other churches of Angers, the principal are St. Serge, an abbey-church of the 12th and 15th centuries, and La Trinit? . The prefecture occupies the buildings of the famous abbey of St. Aubin; in its courtyard are elaborately sculptured arcades of the 11th and 12th centuries, from which period dates the tower, the only survival of the splendid abbey-church. Ruins of the old churches of Toussaint and Notre-Dame du Ronceray are also to be seen. The castle of Angers, an imposing building girt with towers and a moat, dates from the 13th century and is now used as an armoury. The ancient hospital of St. Jean is occupied by an archaeological museum; and the Logis Barrault, a mansion built about 1500, contains the public library, the municipal museum, which has a large collection of pictures and sculptures, and the Mus?e David, containing works by the famous sculptor David d'Angers, who was a native of the town. One of his masterpieces, a bronze statue of Ren? of Anjou, stands close by the castle. The H?tel de Pinc? or d'Anjou is the finest of the stone mansions of Angers; there are also many curious wooden houses of the 15th and 16th centuries. The palais de justice, the Catholic institute, a fine theatre, and a hospital with 1500 beds are the more remarkable of the modern buildings of the town. Angers is the seat of a bishopric, dating from the 3rd century, a prefecture, a court of appeal and a court of assizes. It has a tribunal of first instance, a tribunal of commerce, a board of trade-arbitrators, a chamber of commerce, a branch of the Bank of France and several learned societies. Its educational institutions include ecclesiastical seminaries, a lyc?e, a preparatory school of medicine and pharmacy, a university with free faculties of theology, law, letters and science, a higher school of agriculture, training colleges, a school of arts and handicrafts and a school of fine art. The prosperity of the town is largely due to the great slate-quarries of the vicinity, but the distillation of liqueurs from fruit, cable, rope and thread-making, and the manufacture of boots and shoes, umbrellas and parasols are leading industries. The weaving of sail-cloth and woollen and other fabrics, machine construction, wire-drawing, and manufacture of sparkling wines and preserved fruits are also carried on. The chief articles of commerce, besides slate and manufactured goods, are hemp, early vegetables, fruit, flowers and live-stock.

Angers, capital of the Gallic tribe of the Andecavi, was under the Romans called Juliomagus. During the 9th century it became the seat of the counts of Anjou . It suffered severely from the invasions of the Northmen in 845 and the succeeding years, and of the English in the 12th and 15th centuries; the Huguenots took it in 1585, and the Vendean royalists were repulsed near it in 1793. Till the Revolution, Angers was the seat of a celebrated university founded in the 14th century.

ANGERSTEIN, JOHN JULIUS , London merchant, and patron of the fine arts, was born at St. Petersburg and settled in London about 1749. His collection of paintings, consisting of about forty of the most exquisite specimens of the art, purchased by the British government, on his death, formed the nucleus of the National Gallery.

ANGINA PECTORIS , a term applied to a violent paroxysm of pain, arising almost invariably in connexion with disease of the coronary arteries, a lesion causing progressive degeneration of the heart muscle . An attack of angina pectoris usually comes on with a sudden seizure of pain, felt at first over the region of the heart, but radiating through the chest in various directions, and frequently extending down the left arm. A feeling of constriction and of suffocation accompanies the pain, although there is seldom actual difficulty in breathing. When the attack comes on, as it often does, in the course of some bodily exertion, the sufferer is at once brought to rest, and during the continuance of the paroxysm experiences the most intense agony. The countenance becomes pale, the surface of the body cold, the pulse feeble, and death appears to be imminent, when suddenly the attack subsides and complete relief is obtained. The duration of a paroxysm rarely exceeds two or three minutes, but it may last for a longer period. The attacks are apt to recur on slight exertion, and even in aggravated cases without any such exciting cause. Occasionally the first seizure proves fatal; but more commonly death takes place as the result of repeated attacks. Angina pectoris is extremely rare under middle life, and is much more common in males than in females. It must always be regarded as a disorder of a very serious nature. In the treatment of the paroxysm, nitrite of amyl has now replaced all other remedies. It can be carried by the patient in the form of nitrite of amyl pearls, each pearl containing the dose prescribed by the physician. Kept in this way the drug does not lose strength. As soon as the pain begins the patient crushes a pearl in his handkerchief and holds it to his mouth and nose. The relief given in this way is marvellous and usually takes place within a very few seconds. In the rare cases where this drug does not relieve, hypodermic injections of morphia are used. But on account of the well-known dangers of this drug, it should only be administered by a medical man. To prevent recurrence of the attacks something may be done by scrupulous attention to the general health, and by the avoidance of mental and physical strain. But the most important preventive of all is "bed," of which fourteen days must be enforced on the least premonition of anginal pain.

ANGIOSPERMS. The botanical term "Angiosperm" was coined in the form Angiospermae by Paul Hermann in 1690, as the name of that one of his primary divisions of the plant kingdom, which included flowering plants possessing seeds enclosed in capsules, in contradistinction to his Gymnospermae, or flowering plants with achenial or schizo-carpic fruits--the whole fruit or each of its pieces being here regarded as a seed and naked. The term and its antonym were maintained by Linnaeus with the same sense, but with restricted application, in the names of the orders of his class Didynamia. Its use with any approach to its modern scope only became possible after Robert Brown had established in 1827 the existence of truly naked seeds in the Cycadeae and Coniferae, entitling them to be correctly called Gymnosperms. From that time onwards, so long as these Gymnosperms were, as was usual, reckoned as dicotyledonous flowering plants, the term Angiosperm was used antithetically by botanical writers, but with varying limitation, as a group-name for other dicotyledonous plants. The advent in 1851 of Hofmeister's brilliant discovery of the changes proceeding in the embryo-sac of flowering plants, and his determination of the correct relationships of these with the Cryptogamia, fixed the true position of Gymnosperms as a class distinct from Dicotyledons, and the term Angiosperm then gradually came to be accepted as the suitable designation for the whole of the flowering plants other than Gymnosperms, and as including therefore the classes of Dicotyledons and Monocotyledons. This is the sense in which the term is nowadays received and in which it is used here.

The trend of the evolution of the plant kingdom has been in the direction of the establishment of a vegetation of fixed habit and adapted to the vicissitudes of a life on land, and the Angiosperms are the highest expression of this evolution and constitute the dominant vegetation of the earth's surface at the present epoch. There is no land-area from the poles to the equator, where plant-life is possible, upon which Angiosperms are not found. They occur also abundantly in the shallows of rivers and fresh-water lakes, and in less number in salt lakes and in the sea; such aquatic Angiosperms are not, however, primitive forms, but are derived from immediate land-ancestors. Associated with this diversity of habitat is great variety in general form and manner of growth. The familiar duckweed which covers the surface of a pond consists of a tiny green "thalloid" shoot, one, that is, which shows no distinction of parts--stem and leaf, and a simple root growing vertically downwards into the water. The great forest-tree has a shoot, which in the course perhaps of hundreds of years, has developed a wide-spreading system of trunk and branches, bearing on the ultimate twigs or branchlets innumerable leaves, while beneath the soil a widely-branching root-system covers an area of corresponding extent. Between these two extremes is every conceivable gradation, embracing aquatic and terrestrial herbs, creeping, erect or climbing in habit, shrubs and trees, and representing a much greater variety than is to be found in the other subdivision of seed-plants, the Gymnosperms.

In internal structure also the variety of tissue-formation far exceeds that found in Gymnosperms . The vascular bundles of the stem belong to the collateral type, that is to say, the elements of the wood or xylem and the bast or phloem stand side by side on the same radius. In the larger of the two great groups into which the Angiosperms are divided, the Dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue, known as cambium; by the formation of a layer of cambium between the bundles a complete ring is formed, and a regular periodical increase in thickness results from it by the development of xylem on the inside and phloem on the outside. The soft phloem soon becomes crushed, but the hard wood persists, and forms the great bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth--the so-called annual rings. In the smaller group, the Monocotyledons, the bundles are more numerous in the young stem and scattered through the ground tissue. Moreover they contain no cambium and the stem once formed increases in diameter only in exceptional cases.

In rare cases the main axis is unbranched and ends in a flower, as, for instance, in the tulip, where scale-leaves, forming the underground bulb, green foliage-leaves and coloured floral leaves are borne on one and the same axis. Generally, flowers are formed only on shoots of a higher order, often only on the ultimate branches of a much branched system. A potential branch or bud, either foliage or flower, is formed in the axil of each leaf; sometimes more than one bud arises, as for instance in the walnut, where two or three stand in vertical series above each leaf. Many of the buds remain dormant, or are called to development under exceptional circumstances, such as the destruction of existing branches. For instance, the clipping of a hedge or the lopping of a tree will cause to develop numerous buds which may have been dormant for years. Leaf-buds occasionally arise from the roots, when they are called adventitious; this occurs in many fruit trees, poplars, elms and others. For instance, the young shoots seen springing from the ground around an elm are not seedlings but root-shoots. Frequently, as in many Dicotyledons, the primary root, the original root of the seedling, persists throughout the life of the plant, forming, as often in biennials, a thickened tap-root, as in carrot, or in perennials, a much-branched root system. In many Dicotyledons and most Monocotyledons, the primary root soon perishes, and its place is taken by adventitious roots developed from the stem.

The most characteristic feature of the Angiosperm is the flower, which shows remarkable variety in form and elaboration, and supplies the most trustworthy characters for the distinction of the series and families or natural orders, into which the group is divided. The flower is a shoot which has a special form associated with the special function of ensuring the fertilization of the egg and the development of fruit containing seed. Except where it is terminal it arises, like the leaf-shoot, in the axil of a leaf, which is then known as a bract. Occasionally, as in violet, a flower arises singly in the axil of an ordinary foliage-leaf; it is then termed axillary. Generally, however, the flower-bearing portion of the plant is sharply distinguished from the foliage leaf-bearing or vegetative portion, and forms a more or less elaborate branch-system in which the bracts are small and scale-like. Such a branch-system is called an inflorescence. The primary function of the flower is to bear the spores. These, as in Gymnosperms, are of two kinds, microspores or pollen-grains, borne in the stamens and megaspores, in which the egg-cell is developed, contained in the ovule, which is borne enclosed in the carpel . The flower may consist only of spore-bearing leaves, as in willow, where each flower comprises only a few stamens or two carpels. Usually, however, other leaves are present which are only indirectly concerned with the reproductive process, acting as protective organs for the sporophylls or forming an attractive envelope. These form the perianth and are in one series, when the flower is termed monochlamydeous, or in two series . In the second case the outer series is generally green and leaf-like, its function being to protect the rest of the flower, especially in the bud; while the inner series is generally white or brightly coloured, and more delicate in structure, its function being to attract the particular insect or bird by agency of which pollination is effected. The insect, &c., is attracted by the colour and scent of the flower, and frequently also by honey which is secreted in some part of the flower.

Each stamen generally bears four pollen-sacs which are associated to form the anther, and carried up on a stalk or filament. The development of the microsporangia and the contained spores is closely comparable with that of the microsporangia in Gymnosperms or heterosporous ferns. The pollen is set free by the opening of the anther, generally by means of longitudinal slits, but sometimes by pores, as in the heath family , or by valves, as in the barberry. It is then dropped or carried by some external agent, wind, water or some member of the animal kingdom, on to the receptive surface of the carpel of the same or another flower. The carpel, or aggregate of carpels forming the pistil or gynaeceum, comprises an ovary containing one or more ovules and a receptive surface or stigma; the stigma is sometimes carried up on a style. The mature pollen-grain is, like other spores, a single cell; except in the case of some submerged aquatic plants, it has a double wall, a thin delicate wall of unaltered cellulose, the endospore or intine, and a tough outer cuticularized exospore or extine. The exospore often bears spines or warts, or is variously sculptured, and the character of the markings is often of value for the distinction of genera or higher groups. Germination of the microspore begins before it leaves the pollen-sac. In very few cases has anything representing prothallial development been observed; generally a small cell is cut off, leaving a larger tube-cell. When placed on the stigma, under favourable circumstances, the pollen-grain puts forth a pollen-tube which grows down the tissue of the style to the ovary, and makes its way along the placenta, guided by projections or hairs, to the mouth of an ovule. The nucleus of the tube-cell has meanwhile passed into the tube, as does also the generative nucleus which divides to form two male- or sperm-cells. The male-cells are carried to their destination in the tip of the pollen-tube.

The ovary contains one or more ovules borne on a placenta, which is generally some part of the ovary-wall. The development of the ovule, which represents the macrosporangium, is very similar to the process in Gymnosperms; when mature it consists of one or two coats surrounding the central nucellus, except at the apex where an opening, the micropyle, is left. The nucellus is a cellular tissue enveloping one large cell, the embryo-sac or macrospore. The germination of the macrospore consists in the repeated division of its nucleus to form two groups of four, one group at each end of the embryo-sac. One nucleus from each group, the polar nucleus, passes to the centre of the sac, where the two fuse to form the so-called definitive nucleus. Of the three cells at the micropylar end of the sac, all naked cells , one is the egg-cell or oosphere, the other two, which may be regarded as representing abortive egg-cells , are known as synergidae. The three cells at the opposite end are known as antipodal cells and become invested with a cell-wall. The gametophyte or prothallial generation is thus extremely reduced, consisting of but little more than the male and female sexual cells--the two sperm-cells in the pollen-tube and the egg-cell in the embryo-sac.

At the period of fertilization the embryo-sac lies in close proximity to the opening of the micropyle, into which the pollen-tube has penetrated, the separating cell-wall becomes absorbed, and the male or sperm-cells are ejected into the embryo-sac. Guided by the synergidae one male-cell passes into the oosphere with which it fuses, the two nuclei uniting, while the other fuses with the definitive nucleus, or, as it is also called, the endosperm nucleus. This remarkable double fertilization as it has been called, although only recently discovered, has been proved to take place in widely-separated families, and both in Monocotyledons and Dicotyledons, and there is every probability that, perhaps with variations, it is the normal process in Angiosperms. After impregnation the fertilized oosphere immediately surrounds itself with a cell-wall and becomes the oospore which by a process of growth forms the embryo of the new plant. The endosperm-nucleus divides rapidly to produce a cellular tissue which fills up the interior of the rapidly-growing embryo-sac, and forms a tissue, known as endosperm, in which is stored a supply of nourishment for the use later on of the embryo. It has long been known that after fertilization of the egg has taken place, the formation of endosperm begins from the endosperm nucleus, and this had come to be regarded as the recommencement of the development of a prothallium after a pause following the reinvigorating union of the polar nuclei. This view is still maintained by those who differentiate two acts of fertilization within the embryo-sac, and regard that of the egg by the first male-cell, as the true or generative fertilization, and that of the polar nuclei by the second male gamete as a vegetative fertilization which gives a stimulus to development in correlation with the other. If, on the other hand, the endosperm is the product of an act of fertilization as definite as that giving rise to the embryo itself, we have to recognize that twin-plants are produced within the embryo-sac--one, the embryo, which becomes the angiospermous plant, the other, the endosperm, a short-lived, undifferentiated nurse to assist in the nutrition of the former, even as the subsidiary embryos in a pluri-embryonic Gymnosperm may facilitate the nutrition of the dominant one. If this is so, and the endosperm like the embryo is normally the product of a sexual act, hybridization will give a hybrid endosperm as it does a hybrid embryo, and herein we may have the explanation of the phenomenon of xenia observed in the mixed endosperms of hybrid races of maize and other plants, regarding which it has only been possible hitherto to assert that they were indications of the extension of the influence of the pollen beyond the egg and its product. This would not, however, explain the formation of fruits intermediate in size and colour between those of crossed parents. The signification of the coalescence of the polar nuclei is not explained by these new facts, but it is noteworthy that the second male-cell is said to unite sometimes with the apical polar nucleus, the sister of the egg, before the union of this with the basal polar one. The idea of the endosperm as a second subsidiary plant is no new one; it was suggested long ago in explanation of the coalescence of the polar nuclei, but it was then based on the assumption that these represented male and female cells, an assumption for which there was no evidence and which was inherently improbable. The proof of a coalescence of the second male nucleus with the definitive nucleus gives the conception a more stable basis. The antipodal cells aid more or less in the process of nutrition of the developing embryo, and may undergo multiplication, though they ultimately disintegrate, as do also the synergidae. As in Gymnosperms and other groups an interesting qualitative change is associated with the process of fertilization. The number of chromosomes in the nucleus of the two spores, pollen-grain and embryo-sac, is only half the number found in an ordinary vegetative nucleus; and this reduced number persists in the cells derived from them. The full number is restored in the fusion of the male and female nuclei in the process of fertilization, and remains until the formation of the cells from which the spores are derived in the new generation.

The formation of endosperm starts, as has been stated, from the endosperm nucleus. Its segmentation always begins before that of the egg, and thus there is timely preparation for the nursing of the young embryo. If in its extension to contain the new formations within it the embryo-sac remains narrow, endosperm formation proceeds upon the lines of a cell-division, but in wide embryo-sacs the endosperm is first of all formed as a layer of naked cells around the wall of the sac, and only gradually acquires a pluricellular character, forming a tissue filling the sac. The function of the endosperm is primarily that of nourishing the embryo, and its basal position in the embryo-sac places it favourably for the absorption of food material entering the ovule. Its duration varies with the precocity of the embryo. It may be wholly absorbed by the progressive growth of the embryo within the embryo-sac, or it may persist as a definite and more or less conspicuous constituent of the seed. When it persists as a massive element of the seed its nutritive function is usually apparent, for there is accumulated within its cells reserve-food, and according to the dominant substance it is starchy, oily, or rich in cellulose, mucilage or proteid. In cases where the embryo has stored reserve food within itself and thus provided for self-nutrition, such endosperm as remains in the seed may take on other functions, for instance, that of water-absorption.

Their fortuitous dissemination does not always bring seeds upon a suitable nidus for germination, the primary essential of which is a sufficiency of moisture, and the duration of vitality of the embryo is a point of interest. Some seeds retain vitality for a period of many years, though there is no warrant for the popular notion that genuine "mummy wheat" will germinate; on the other hand some seeds lose vitality in little more than a year. Further, the older the seed the more slow as a general rule will germination be in starting, but there are notable exceptions. This pause, often of so long duration, in the growth of the embryo between the time of its perfect development within the seed and the moment of germination, is one of the remarkable and distinctive features of the life of Spermatophytes. The aim of germination is the fixing of the embryo in the soil, effected usually by means of the root, which is the first part of the embryo to appear, in preparation for the elongation of the epicotyledonary portion of the shoot, and there is infinite variety in the details of the process. In albuminous Dicotyledons the cotyledons act as the absorbents of the reserve-food of the seed and are commonly brought above ground , either withdrawn from the seed-coat or carrying it upon them, and then they serve as the first green organs of the plant. The part of the stem below the cotyledons commonly plays the greater part in bringing this about. Exalbuminous Dicotyledons usually store reserve-food in their cotyledons, which may in germination remain below ground . In albuminous Monocotyledons the cotyledon itself, probably in consequence of its terminal position, is commonly the agent by which the embryo is thrust out of the seed, and it may function solely as a feeder, its extremity developing as a sucker through which the endosperm is absorbed, or it may become the first green organ, the terminal sucker dropping off with the seed-coat when the endosperm is exhausted. Exalbuminous Monocotyledons are either hydrophytes or strongly hygrophilous plants and have often peculiar features in germination.

Distribution by seed appears to satisfy so well the requirements of Angiosperms that distribution by vegetative buds is only an occasional process. At the same time every bud on a shoot has the capacity to form a new plant if placed in suitable conditions, as the horticultural practice of propagation by cuttings shows; in nature we see plants spreading by the rooting of their shoots, and buds we know may be freely formed not only on stems but on leaves and on roots. Where detachable buds are produced, which can be transported through the air to a distance, each of them is an incipient shoot which may have a root, and there is always reserve-food stored in some part of it. In essentials such a bud resembles a seed. A relation between such vegetative distribution buds and production of flower is usually marked. Where there is free formation of buds there is little flower and commonly no seed, and the converse is also the case. Viviparous plants are an illustration of substitution of vegetative buds for flower.

The position of Angiosperms as the highest plant-group is unassailable, but of the point or points of their origin from the general stem of the plant kingdom, and of the path or paths of their evolution, we can as yet say little.

Until well on in the Mesozoic period geological history tells us nothing about Angiosperms, and then only by their vegetative organs. We readily recognize in them now-a-days the natural classes of Dicotyledons and Monocotyledons distinguished alike in vegetative and in reproductive construction, yet showing remarkable parallel sequences in development; and we see that the Dicotyledons are the more advanced and show the greater capacity for further progressive evolution. But there is no sound basis for the assumption that the Dicotyledons are derived from Monocotyledons; indeed, the palaeontological evidence seems to point to the Dicotyledons being the older. This, however, does not entitle us to assume the origin of Monocotyledons from Dicotyledons, although there is manifestly a temptation to connect helobic forms of the former with ranal ones of the latter. There is no doubt that the phylum of Angiosperms has not sprung from that of Gymnosperms.

Within each class the flower-characters as the essential feature of Angiosperms supply the clue to phylogeny, but the uncertainty regarding the construction of the primitive angiospermous flower gives a fundamental point of divergence in attempts to construct progressive sequences of the families. Simplicity of flower-structure has appeared to some to be always primitive, whilst by others it has been taken to be always derived. There is, however, abundant evidence that it may have the one or the other character in different cases. Apart from this, botanists are generally agreed that the concrescence of parts of the flower-whorls--in the gynaeceum as the seed-covering, and in the corolla as the seat of attraction, more than in the androecium and the calyx--is an indication of advance, as is also the concrescence that gives the condition of epigyny. Dorsiventrality is also clearly derived from radial construction, and anatropy of the ovule has followed atropy. We should expect the albuminous state of the seed to be an antecedent one to the exalbuminous condition, and the recent discoveries in fertilization tend to confirm this view. Amongst Dicotyledons the gamopetalous forms are admitted to be the highest development and a dominant one of our epoch. Advance has been along two lines, markedly in relation to insect-pollination, one of which has culminated in the hypogynous epipetalous bicarpellate forms with dorsiventral often large and loosely arranged flowers such as occur in Scrophulariaceae, and the other in the epigynous bicarpellate small-flowered families of which the Compositae represent the most elaborate type. In the polypetalous forms progression from hypogyny to epigyny is generally recognized, and where dorsiventrality with insect-pollination has been established, a dominant group has been developed as in the Leguminosae. The starting-point of the class, however, and the position within it of apetalous families with frequently unisexual flowers, have provoked much discussion. In Monocotyledons a similar advance from hypogyny to epigyny is observed, and from the dorsiventral to the radial type of flower. In this connexion it is noteworthy that so many of the higher forms are adapted as bulbous geophytes, or as aerophytes to special xerophilous conditions. The Gramineae offer a prominent example of a dominant self-pollinated or wind-pollinated family, and this may find explanation in a multiplicity of factors.

DICOTYLEDONS.

Polypetalae: Thalamiflorae. Disciflorae. Calyciflorae.

Gamopetalae: Inferae. Heteromerae. Bicarpellatae.

Monochlamydeae in eight series. Monocotyledons in seven series.

Of the Polypetalae, series 1, Thalamiflorae, is characterized by hypogynous petals and stamens, and contains 34 orders distributed in 6 larger groups or cohorts. Series 2, Disciflorae, takes its name from a development of the floral axis which forms a ring or cushion at the base of the ovary or is broken up into glands; the ovary is superior. It contains 23 orders in 4 cohorts. Series 3, Calyciflorae, has petals and stamens perigynous, or sometimes superior. It contains 27 orders in 5 cohorts.

Of the Gamopetalae, series 1, Inferae, has an interior ovary and stamens usually as many as the corolla-lobes. It contains 9 orders in 3 cohorts. Series 2, Heteromerae, has generally a superior ovary, stamens as many as the corolla-lobes or more, and more than two carpels. It contains 12 orders in 3 cohorts. Series 3, Bicarpellatae, has generally a superior ovary and usually two carpels. It contains 24 orders in 4 cohorts.

The eight series of Monochlamydeae, containing 36 orders, form groups characterized mainly by differences in the ovary and ovules, and are now recognized as of unequal value.

The seven series of Monocotyledons represent a sequence beginning with the most complicated epigynous orders, such as Orchideae and Scitamineae, and passing through the petaloid hypogynous orders of which Liliaceae is the representative to Juncaceae and the palms where the perianth loses its petaloid character and thence to the Aroids, screw-pines and others where it is more or less aborted . Series 6, Apocarpeae, is characterized by 5 carpels, and in the last series Glumaceae, great simplification in the flower is associated with a grass-like habit.

As an attempt at a phylogenetic arrangement, Engler's system is now preferred by many botanists. More recently a startling novelty in the way of system has been produced by van Tieghem, as follows:

The most remarkable feature here is the class of Liorhizal Dicotyledons, which includes only the families of Nymphaeaceae and Gramineae. It is based upon the fact that the histological differentiation of the epidermis of their root is that generally characteristic of Monocotyledons, whilst they have two cotyledons--the old view of the epiblast as a second cotyledon in Gramineae being adopted. But the presence of a second cotyledon in grasses is extremely doubtful, and though there may be ground for reconsidering the position of Nymphaeaceae, their association with the grasses as a distinct class is not warranted by a comparative examination of the members of the two orders. Ovular characters determine the grouping in the Dicotyledons, van Tieghem supporting the view that the integument, the outer if there be two, is the lamina of a leaf of which the funicle is the petiole, whilst the nucellus is an outgrowth of this leaf, and the inner integument, if present, an indusium. The Insemineae include forms in which the nucellus is not developed, and therefore there can be no seed. The plants included are, however, mainly well-established parasites, and the absence of nucellus is only one of those characters of reduction to which parasites are liable. Even if we admit van Tieghem's interpretation of the integuments to be correct, the diagnostic mark of his unitegminous and bitegminous groups is simply that of the absence or presence of an indusium, not a character of great value elsewhere, and, as we know, the number of the ovular coats is inconstant within the same family. At the same time the groups based upon the integuments are of much the same extent as the Polypetalae and Gamopetalae of other systems. We do not yet know the significance of this correlation, which, however, is not an invariable one, between number of integuments and union of petals.

Within the last few years Prof. John Coulter and Dr. C.J. Chamberlain of Chicago University have given a valuable general account of the morphology of Angiosperms as far as concerns the flower, and the series of events which ends in the formation of the seed .

ANGKOR, an assemblage of ruins in Cambodia, the relic of the ancient Khmer civilization. They are situated in forests to the north of the Great Lake , the most conspicuous of the remains being the town of Angkor-Thom and the temple of Angkor-Vat, both of which lie on the right bank of the river Siem-Reap, a tributary of Tonle-Sap. Other remains of the same form and character lie scattered about the vicinity on both banks of the river, which is crossed by an ancient stone bridge.

The vestiges of the royal palace, which stood within an enclosure containing also the pyramidal religious structure known as the Phimeanakas. To the east of this enclosure there extends a terrace decorated with magnificent reliefs.

The temple of Bayon, a square enclosure formed by galleries with colonnades, within which is another and more elaborate system of galleries, rectangular in arrangement and enclosing a cruciform structure, at the centre of which rises a huge tower with a circular base. Fifty towers, decorated with quadruple faces of Brahma, are built at intervals upon the galleries, the whole temple ranking as perhaps the most remarkable of the Khmer remains.

Angkor-Vat, the best preserved example of Khmer architecture, lies less than a mile to the south of the royal city, within a rectangular park surrounded by a moat, the outer perimeter of which measures 6060 yds. On the west side of the park a paved causeway, leading over the moat and under a magnificent portico, extends for a distance of a quarter of a mile to the chief entrance of the main building. The temple was originally devoted to the worship of Brahma, but afterwards to that of Buddha; its construction is assigned by Aymonier to the first half of the 12th century A.D. It consists of three stages, connected by numerous exterior staircases and decreasing in dimensions as they rise, culminating in the sanctuary, a great central tower pyramidal in form. Towers also surmount the angles of the terraces of the two upper stages. Three galleries with vaulting supported on columns lead from the three western portals to the second stage. They are connected by a transverse gallery, thus forming four square basins. Khmer decoration, profuse but harmonious, consists chiefly in the representation of gods, men and animals, which are displayed on every flat surface. Combats and legendary episodes are often depicted; floral decoration is reserved chiefly for borders, mouldings and capitals. Sandstone of various colours was the chief material employed by the Khmers; limonite was also used. The stone was cut into huge blocks which are fitted together with great accuracy without the use of cement.

Lord Anglesey married Elizabeth, daughter and co-heiress of Sir James Altham of Oxey, Hertfordshire, by whom, besides other children, he had James, who succeeded him, Altham, created Baron Altham, and Richard, afterwards 3rd Baron Altham. His descendant Richard, the 6th earl , left a son Arthur, whose legitimacy was doubted, and the peerage became extinct. He was summoned to the Irish House of Peers as Viscount Valentia, but was denied his writ to the parliament of Great Britain by a majority of one vote. He was created in 1793 earl of Mountnorris in the peerage of Ireland. All the male descendants of the 1st earl of Anglesey became extinct in the person of George, 2nd earl of Mountnorris, in 1844, when the titles of Viscount Valentia and Baron Mountnorris passed to his cousin Arthur Annesley , who thus became 10th Viscount Valentia, being descended from the 1st Viscount Valentia the father of the 1st earl of Anglesey in the Annesley family. The 1st viscount was also the ancestor of the Earls Annesley in the Irish peerage.

The marquess had a large family by each of his two wives, two sons and six daughters by the first and six sons and four daughters by the second. His eldest son, Henry, succeeded him in the marquessate; but the title passed rapidly in succession to the 3rd, 4th and 5th marquesses. The latter, whose extravagances were notorious, died in 1905, when the title passed to his cousin.

Other members of the Paget family distinguished themselves in the army and the navy. Of the first marquess's brothers one, SIR CHARLES PAGET , rose to the rank of vice-admiral in the Royal Navy; another, General SIR EDWARD PAGET , won great distinction by his skilful and resolute handling of a division at Corunna, and from 1822 to 1825 was commander-in-chief in India. One of the marquess's sons by his second marriage, LORD CLARENCE EDWARD PAGET , became an admiral; another, LORD GEORGE AUGUSTUS FREDERICK PAGET , led the 4th Light Dragoons in the charge of the Light Brigade at Balaklava, and subsequently commanded the brigade, and, for a short time, the cavalry division in the Crimea. In 1865 he was made inspector-general of cavalry, in 1871 lieutenant-general and K.C.B., and in 1877 full general. His Crimean journals were published in 1881.

ANGLESEY, or ANGLESEA, an insular northern county of Wales. Its area is 176,630 acres or about 276 sq. m. Anglesey, in the see of Bangor, is separated from the mainland by the Menai Straits , over which were thrown Telford's suspension bridge, in 1826, and the Stephenson tubular railway bridge in 1850. The county is flat, with slight risings such as Parys, Cadair Mynachdy and Holyhead Mountain. There are a few lakes, such as Cors cerrig y daran, but rising water is generally scarce. The climate is humid, the land poor for the most part compared with its old state of fertility, and there are few industries.

As regards geology, the younger strata in Anglesey rest upon a foundation of very old pre-Cambrian rocks which appear at the surface in three areas:-- a western region including Holyhead and Llanfaethlu, a central area about Aberffraw and Trefdraeth, and an eastern region which includes Newborough, Caerwen and Pentraeth. These pre-Cambrian rocks are schists and slates, often much contorted and disturbed. The general line of strike of the formations in the island is from N.E. to S.W. A belt of granitic rocks lies immediately north-west of the central pre-Cambrian mass, reaching from Llanfaelog near the coast to the vicinity of Llanerchymedd. Between this granite and the pre-Cambrian of Holyhead is a narrow tract of Ordovician slates and grits with Llandovery beds in places; this tract spreads out in the N. of the island between Dulas Bay and Carmel Point. A small patch of Ordovician strata lies on the northern side of Beaumaris. In parts, these Ordovician rocks are much folded, crushed and metamorphosed, and they are associated with schists and altered volcanic rocks which are probably pre-Cambrian. Between the eastern and central pre-Cambrian masses carboniferous rocks are found. The carboniferous limestone occupies a broad area S. of Ligwy Bay and Pentraeth, and sends a narrow spur in a south-westerly direction by Llangefni to Malldraeth sands. The limestone is underlain on the N.W. by a red basement conglomerate and yellow sandstone . Limestone occurs again on the N. coast about Llanfihangel and Llangoed; and in the S.W. round Llanidan on the border of the Menai Strait. Puffin Island is made of carboniferous limestone. Malldraeth Marsh is occupied by coal measures, and a small patch of the same formation appears near Tall-y-foel Ferry on the Menai Straits. A patch of granitic and felsitic rocks form Parys Mountain, where copper and iron ochre have been worked. Serpentine is found near Llanfaerynneubwll and upon the opposite shore in Holyhead. There are abundant evidences of glaciation, and much boulder clay and drift sand covers the older rocks. Patches of blown sand occur on the S.W. coast.

The London & North-Western railway crosses Anglesey from Llanfairpwllgwyngyll to Gaerwen and Holyhead , also from Gaerwen to Amlwch. The staple of the island is farming, the chief crops being turnips, oats, potatoes, with flax in the centre. Copper , lead, silver, marble, asbestos, lime and sandstone, marl, zinc and coal have all been worked in Anglesey, coal especially at Malldraeth and Trefdraeth. The population of the county in 1901 was 50,606. There is no parliamentary borough, but one member is returned for the county. It is in the north-western circuit, and assizes are held at Beaumaris, the only municipal borough . Amlwch , Holyhead , Llangefni and Menai Bridge are urban districts. There are six hundreds and seventy-eight parishes.

Anglesite is a mineral of secondary origin, having been formed by the oxidation of galena in the upper parts of mineral lodes where these have been affected by weathering processes. At Monteponi the crystals encrust cavities in glistening granular galena; and from Leadhills, in Scotland, pseudomorphs of anglesite after galena are known. At most localities it is found as isolated crystals in the lead-bearing lodes, but at some places, in Australia and Mexico, it occurs as large masses, and is then mined as an ore of lead, of which the pure mineral contains 68%.

ANGLICAN COMMUNION, the name used to denote that great branch of the Christian Church consisting of the various churches in communion with the Church of England. The necessity for such a phrase as "Anglican Communion," first used in the 19th century, marked at once the immense development of the Anglican Church in modern times and the change which has taken place in the traditional conceptions of its character and sphere. The Church of England itself is the subject of a separate article ; and it is not without significance that for more than two centuries after the Reformation the history of Anglicanism is practically confined to its developments within the limits of the British Isles. Even in Ireland, where it was for over three centuries the established religion, and in Scotland, where it early gave way to the dominant Presbyterianism, its religious was long overshadowed by its political significance. The Church, in fact, while still claiming to be Catholic in its creeds and in its religious practice, had ceased to be Catholic in its institutional conception, which was now bound up with a particular state and also with a particular conception of that state. To the native Irishman and the Scotsman, as indeed to most Englishmen, the Anglican Church was one of the main buttresses of the supremacy of the English crown and nation. This conception of the relations of church and state was hardly favourable to missionary zeal; and in the age succeeding the Reformation there was no disposition on the part of the English Church to emulate the wonderful activity of the Jesuits, which, in the 16th and 17th centuries, brought to the Church of Rome in countries beyond the ocean compensation for what she had lost in Europe through the Protestant reformation. Even when English churchmen passed beyond the seas, they carried with them their creed, but not their ecclesiastical organization. Prejudice and real or imaginary legal obstacles stood in the way of the erection of episcopal sees in the colonies; and though in the 17th century Archbishop Laud had attempted to obtain a bishop for Virginia, up to the time of the American revolution the churchmen of the colonies had to make the best of the legal fiction that their spiritual needs were looked after by the bishop of London, who occasionally sent commissaries to visit them and ordained candidates for the ministry sent to England for the purpose.

This act had a significance beyond the fact that it established in the United States of America a flourishing church, which, while completely loyal to its own country, is bound by special ties to the religious life of England. It marked the emergence of the Church of England from that insularity to which what may be called the territorial principles of the Reformation had condemned her. The change was slow, and it is not yet by any means complete.

Since the Church of England, whatever her attitude towards the traditional Catholic doctrines, never disputed the validity of Catholic orders whether Roman or Orthodox, nor the jurisdiction of Catholic bishops in foreign countries, the expansion of the Anglican Church has been in no sense conceived as a Protestant aggressive movement against Rome. Occasional exceptions, such as the consecration by Archbishop Plunket of Dublin of a bishop for the reformed church in Spain, raised so strong a protest as to prove the rule. In the main, then, the expansion of the Anglican Church has followed that of the British empire, or, as in America, of its daughter states; its claim, so far as rights of jurisdiction are concerned, is to be the Church of England and the English race, while recognizing its special duties towards the non-Christian populations subject to the empire or brought within the reach of its influence. As against the Church of Rome, with its system of rigid centralization, the Anglican Church represents the principle of local autonomy, which it holds to be once more primitive and more catholic. In this respect the Anglican communion has developed on the lines defined in her articles at the Reformation; but, though in principle there is no great difference between a church defined by national, and a church defined by racial boundaries, there is an immense difference in effect, especially when the race--as in the case of the English--is itself ecumenical.

The realization of what may be called this catholic mission of the English church, in the extension of its organization to the colonies, was but a slow process.

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